Braby, M.F., Espeland, M., Müller, C.J., Eastwood, R., Lohman, D.J., Kawahara, A.Y., Maunsell, S.C. and Pierce, N.E. (2020), Molecular phylogeny of the tribe Candalidini (Lepidoptera: Lycaenidae): systematics, diversification and evolutionary history. Syst Entomol, 45: 703-722. (Linkki)