https://doi.org/10.1017/S0024282917000615 and https://doi.org/10.1017/S0024282919000203

The first article is the genetic work, but the second article sorts out the nomenclature much better. What the first article called "Thamnolia subuliformis" is really Thamnolia vermicularis subsp. vermicularis, what it called "Thamnolia vermicularis" is really Thamnolia vermicularis subsp. taurica and their new species was demoted to a subspecies. There is very little genetic difference between these taxa, and only taurica and tundrae can be distinguished from each other by chemistry (but they are not found in the same areas, anyway). Both taurica and tundrae cannot be separated from vermicularis by any means but sequencing, as vermicularis can have the same chemistry as either, depending.